unfavorable sites for Plant growth, and an R- Five traits that are coordinated across species are axis (Ruderal species) reflects adaptation to leaf mass per area (LMA), leaf life-span, leaf N disturbance.. unfavorable sites for Plant growth, and an R- Five traits that are coordinated across species are axis (Ruderal species) reflects adaptation to leaf mass per area (LMA), leaf life-span, leaf N disturbance.